Is the 9+2 pattern of flagellar and ciliary axonemes an efficient arrangement for generating planar bending?

Journal of mechanochemistry & cell motility

PubMedID: 753898

Brokaw CJ. Is the 9+2 pattern of flagellar and ciliary axonemes an efficient arrangement for generating planar bending?. J Mechanochem Cell Motil. 1977;4(2):101-11.
A circlet of 9 interacting microtubular doublets generates less bending moment at low bending rates, but allows bending moment to be generated at higher bending rates, and has an energy efficiency at least 92% as great as an arrangement with all the interactions between microtubular doublets in a plane parallel to the bending plane. The axonemal arrangement has no significant effect on the shape of the normalized relationship between active shear moment and shear rate.